29Morphology of Angiosperm

▢ Introduction:

Table 1: Morphology Basics

Term / Point
Meaning
Morphology
Study of form, structure, size, colour, development and relative position of organism parts
Anthology
Study of flowering plants / flowers
Angiosperms
Flowering plants
Phanerogamic
Flowering / visible reproductive organs
Spermatophytic
Seed-bearing plants
Sporophytic
Main plant body diploid / 2n
First flowering plants
Appeared in Jurassic period
▢ Comparative Morphology:

Table 1: Root, Stem, Leaf, Inflorescence and Flower

Part
Arrangement
Stalk / Axis
Special term
Root
No node and internode
Root axis
Descending axis
Stem
Nodes and internodes present
Stem axis
Ascending axis
Leaf
Phyllotaxy = arrangement of mature leaves on stem/branches
Petiole
Prefoliation = arrangement of young leaves in bud
Inflorescence
Arrangement and distribution of flowers on shoot system
Peduncle
Anthotaxy = arrangement of flowers in inflorescence
Flower
Floral whorls on condensed axis
Pedicel
Thalamus / torus = swollen end of pedicel
▢ Phyllotaxy and Prefoliation:
❖ Phyllotaxy:
◉ Definition: Arrangement of mature leaves on stem or branches

Table 1: Types of Phyllotaxy

Type
Feature
Alternate / Spiral
Single leaf from each node
Opposite
Pair of leaves arise from node opposite to each other
Opposite superposed
Leaves of successive nodes in same plane → two rows
Opposite decussate
Opposite leaves of one node at right angle to adjacent node
Whorled / Verticillate
Three or more leaves from a single node
❖ Prefoliation:
◉ Definition: Arrangement of immature / young leaves in bud

Table 1: Prefoliation

Type
Meaning
Example
Ptyxis
Arrangement of individual immature leaf in bud
Circinate
Circinate ptyxis
Young leaf coiled like watch spring
Fern, Cycas, banana
Vernation
Arrangement of immature leaves with respect to each other in bud
Leaf bud
Aestivation
Arrangement of floral parts in floral bud
Sepals / petals
📚
ROOT
▢ Introduction:

Table 1: Root Characters

Feature
Point
Position
Usually underground
Colour
Non-green
Direction
Descending part of plant
Geotropism
Positively geotropic
Phototropism
Negatively phototropic
First structure in germination
Root / radicle
Main root
Primary root / tap root from radicle
Lateral roots
Arise from pericycle; endogenous
Adventitious root
Root from any part other than radicle
Tissue culture root
Arises from callus
❖ Apogeotropic Roots:
  • •
    Coralloid roots of Cycas
  • •
    Pneumatophores of mangrove plants
❖ Functions:
  • •
    Fixation
  • •
    Absorption
  • •
    Conduction
  • •
    Assimilation in some roots
  • •
    Reproduction in some roots
❖ Absent in Root:
  • •
    Nodes and internodes
  • •
    Buds generally absent
  • •
    Leaves
  • •
    Flowers
▢ Special Plants:

Table 1: Plantless Roots and Rootless Plants

Category
Examples
Plantless roots
Podostemon, Arceuthobium, Rafflesia, Sapria, Monotropa
Rootless plants
Myriophyllum, Balanophora, Utricularia, Ceratophyllum, Cuscuta, Wolffia
📖
Tap root modifications
▢ Fleshy Tap Roots:

Table 1: Storage Tap Roots

Type
Shape
Example
Fusiform
Spindle-shaped; tapering at both ends
Raphanus sativus / radish
Napiform
Top-shaped; swollen base, abruptly tapering apex
Brassica rapa / turnip
Conical
Cone-shaped; broad base, tapering apex
Daucus carota / carrot
Tuberous
Irregular swelling
Mirabilis jalapa / four o'clock plant
▢ Other Tap Root Modifications:

Table 1: Tap Root Modifications

Type
Feature
Examples
Nodulated / tuberculated root
Root nodules in legumes; nitrogen fixation by Rhizobium; leghaemoglobin protects nitrogenase and supplies O₂
Leguminous plants
Pneumatophores / respiratory roots
Negatively geotropic roots with pneumatophores / breathing pores
Rhizophora, Avicennia
Buttress roots
Laterally compressed horizontal roots at base of trunk for support
Ficus religiosa, Ficus elastica, Bombax ceiba
Mycorrhizal roots
Root cap and root hairs absent; fungal hyphae cover root; phosphorus absorption
Pinus, Betula, orchids
Reproductive tap root
Develops adventitious buds for vegetative propagation
Dalbergia
❖ Pneumatothodes: Minute pores on exposed pneumatophores; anatomically and morphologically similar to lenticels
📖
Adventitious root modifications
▢ Fleshy Adventitious Roots:

Table 1: Storage Adventitious Roots

Type
Swelling
Examples
Annulated root
Ring-like swelling
Cephaelis, Psychotria / ipecac
Nodulose root
Swollen at tip
Curcuma amada / mango ginger, Curcuma domestica / turmeric
Moniliform / beaded root
Swelling at intervals
Momordica charantia / bitter gourd
Tuberous root / single root tuber
Single, no definite shape
Ipomoea batatas / sweet potato
Fasciculated root
Clustered roots with definite shape
Asparagus, Dahlia
Palmate root
Branched root like palm
Orchis
▢ Support and Special Adventitious Roots:

Table 1: Adventitious Root Modifications

Type
Feature
Examples
Pillar root / prop root
Arises from large branches; thick woody root; support + hygroscopic absorption; lenticels present
Ficus benghalensis, Rhizophora
Stilt root
Arises obliquely from lower nodes; supports plant
Sugarcane, maize, screw pine
Climbing / clinging root
Adventitious roots from node/internode for fixation
Pothos, Piper betle, ivy, vanilla, orchid
Epiphytic root
Clinging root for fixation; hygroscopic root with velamen absorbs moisture
Orchids
Haustorial / parasitic / sucking root
Absorbs nutrition from host
Cuscuta, Viscum
Photosynthetic / assimilatory root
Green roots perform photosynthesis
Trapa, Tinospora, Taeniophyllum
Floating root
Leafy floating roots for balance
Pistia, Eichhornia, Jussiaea
Root thorns
Roots arise from stem and become protective thorns
Pothos, Acanthorhiza
Reproductive root
Adventitious roots produce buds
Dahlia, sweet potato, Trichosanthes dioica
Haptera
Root-like structure from leaf margin; attaches plant to substratum
Podostemon
Contractile root
Arises from underground stem; pulls organ into soil
Crocus, onion, Freesia
Leaf root / foliar root / epiphyllous root
Leaf modified into root for balancing
Salvinia
Nest root
Adventitious roots forming nest-like structure
Dischidia
📚
STEM
▢ Introduction:

Table 1: Stem Characters

Feature
Point
Direction
Ascending part of plant
Origin
Plumule of embryo
Hair
Stem hairs commonly multicellular
Differentiation
Nodes and internodes present
Branches
Produced exogenously from surface meristem in cortex
Structures borne
Branches, leaves, flowers
▢ Aerial Erect Stem:

Table 1: Aerial Erect Stem Types

Type
Feature
Examples
Culm
Distinct nodes and internodes; internode hollow or spongy; node solid
Grass family, sugarcane, maize
Excurrent
Extreme apical dominance; lower branches longer than upper
Most conifers
Deliquescent
Dome / umbrella-shaped crown; branches grow equally in all directions
Mango, peepal
Pseudostem
Aerial unbranched trunk-like structure formed by compact leaf sheaths; true stem underground rhizome
Banana, onion, ginger
Caudex
Unbranched main trunk with tuft of leaves at crown; trunk marked by leaf scars
Phoenix, coconut, Carica, Cycas
▢ Aerial Weak Stem:
❖ Twiners vs Climbers:

Table 1: Twiners vs Climbers

Type
Feature
Examples
Twiner
Main stem coils around support; no special attachment organ; coiling direction genetically determined
Bean, Convolvulus, Clitoria, Dioscorea
Climber
Uses accessory organs for climbing
Root climbers, tendril climbers, scramblers
Sinistrose twiner
Left-handed / anticlockwise coiling
—
Dextrose twiner
Right-handed / clockwise coiling
—
❖ Climbers:

Table 1: Types of Climbers

Type
Modification
Examples
Root climber
Adventitious roots for climbing
Pothos, Piper, betel
Stem tendril climber
Axillary bud → Passiflora; extra-axillary bud → Cucurbita; branch tip → grape
Passiflora, Cucurbita, Vitis vinifera
Leaf tendril climber
Whole leaf / leaflet / leaf tip / petiole / stipule modified
Lathyrus aphaca, pea, sweet pea, Gloriosa, Nepenthes, Smilax
Scrambler / rambler
Uses hooks / pointed structures for climbing
Stragglers
◉ Haptotropism: Coiling of tendril around support due to contact
❖ Thorns, Spines and Prickles:

Table 1: Thorn vs Spine vs Prickle

Structure
Modification / Origin
Examples
Thorn
Modified stem / axillary bud
Duranta, Bougainvillea, citrus
Spine
Modified leaf; bud present in axil
Aloe vera, Opuntia, Asparagus
Prickle
Superficial outgrowth; may arise from any plant part
Climbing rose
❖ Lianas:
  • •
    Large woody climbers and twiners of tropical rain forests
  • •
    Gnetum
  • •
    Bauhinia
  • •
    Bignonia
  • •
    Hiptage
▢ Sub-aerial Stem:
❖ Introduction:
  • •
    Prostrate weak stem modification
  • •
    Produced for vegetative propagation
  • •
    Arises from axil of lower leaves
❖ Types:

Table 1: Sub-aerial Stem Modifications

Type
Feature
Examples
Trailer
Stem spreads horizontally without adventitious roots; fixed at one point by tap root; no daughter plants
Convolvulus, Oxalis, Portulaca, Tridax, Euphorbia prostrata
Runner
Horizontal stem with adventitious roots at nodes/internodes
Oxalis, Cynodon, Lippia, grasses, Centella
Offset
One-internode long, thick, short runner
Pistia, Eichhornia, Agave
Sucker
Obliquely growing running stem
Banana, chrysanthemum, mint
Stolon
Slender lateral branch; on touching soil forms leaves above and roots below
Strawberry, jasmine, blackberry
Sobole
Underground running stem
—
❖ Trailer Types:

Table 1: Trailers

Type
Feature
Examples
Procumbent
Flat branch on ground
Convolvulus, Oxalis
Decumbent
Initially vertical then lies on ground
Portulaca, Tridax
Diffused
Branches spread in all directions
Euphorbia prostrata
▢ Underground Stem:
❖ Introduction:
  • •
    Subterranean stem
  • •
    Plants with underground stem = acaulescent
  • •
    Storage and perennation
  • •
    Vegetative propagation common

Table 1: Underground Stem Modifications

Type
Feature
Examples
Bulb
Condensed disc-like stem with fleshy leaf bases; adventitious roots below
Onion, garlic, lily
Corm
Underground vertical, spherical/globular, unbranched stem; cataphylls; circular nodes
Colocasia, Amorphophallus, Colchicum autumnale
Rhizome
Reduced food-storing stem with cataphylls, nodes, internodes, adventitious roots; diageotropic
Banana, ginger, turmeric, canna
Tuber
Swollen underground branch from axil of lower leaf; nodes arranged spirally; eyes present
Potato
❖ Bulb Types:

Table 1: Bulb Types

Type
Feature
Example
Tunicated bulb
Covered by tunic
Onion, garlic
Scaly / loose bulb
Tunic absent
Lily
❖ Tuber Eye: Each eye of potato has 3 buds → 1 axillary + 2 accessory buds
❖ Largest Corm: Amorphophallus
▢ Stem Modifications:

Table 1: Special Stem Modifications

Type
Feature
Examples
Phylloclade
  1. Green, photosynthetic, succulent stem of indefinite growth
  2. leaves caducous
  3. xerophytic adaptation
  1. Opuntia
  2. Image 1
  3. Euphorbia
  4. Image 1
  5. Muehlenbeckia
  6. Casuarina
Cladode / Cladophyll
  1. Photosynthetic stem of definite growth
  2. usually one/two internodes long
  1. Asparagus
  2. Image 1
  3. Ruscus
  4. Lemna
Thalamus
Condensed axis of flower; edible part of apple
Apple
❖ Mnemonic:
  • •
    Phylloclade → POEM
  • •
    Cladode → CAR
📚
LEAF
▢ Introduction:

Table 1: Leaf Characters

Feature
Point
Definition
Green, exogenous, lateral outgrowth borne on node of stem/branch
Growth
Limited; by intercalary meristem
Major functions
Photosynthesis, gaseous exchange, transpiration
Prophyll
First leaf of plant from shoot meristem as leaf primordium
Phyllome
All types of leaves present in a plant
Single-leaf plant
Monophyllaea
Two-leaf plant
Welwitschia
Absorbing leaf
Lamina of insectivorous plant
Dicot leaf
Dorsiventral + horizontal
Monocot leaf
Isobilateral + vertical
❖ Leaf Surfaces:

Table 1: Leaf Surfaces

Surface
Synonym
Direction
Abaxial surface
Dorsal / lower surface
Away from stem axis
Adaxial surface
Ventral / upper surface
Toward stem axis
📖
Leaf classification
▢ Based on Insertion:

Table 1: Leaf Types by Insertion

Type
Feature
Examples
Radical
Leaves arise above root system from condensed shoot; internodes short
Radish, carrot, turnip
Cauline
Leaves borne on main stem / inflorescence stalk
Mustard, maize, sugarcane
Ramal
Leaves on nodes of stem branches
Mango, peepal
▢ Based on Life Span:

Table 1: Leaf Types by Life Span

Type
Feature
Examples
Caducous
Fall soon after appearance
Opuntia
Deciduous
Fall simultaneously at end of growing season
Acacia catechu, Dalbergia sissoo, mulberry, Populus
Persistent
Live more than one season
Evergreen trees, Pinus, Ficus, mango
▢ Based on Origin and Function:

Table 1: Leaf Types by Origin and Function

Type
Feature
Examples
Foliage leaf
Normal green photosynthetic leaf
Most plants
Reproductive leaf
Produces adventitious roots/shoots for regeneration
Begonia, Bryophyllum
Cotyledonary leaf
Derived from cotyledon; stores food and nourishes seedling
Cucumber, tamarind, castor, bean
Sporophyll
Spore-bearing leaf
Fern frond
Trophophyll
Vegetative / non-spore-bearing leaf
—
Cataphyll
Protective leaf on underground stem
Rhizome, corm
Scale leaf
Reduced leaf found in cladode
Pinus
Hypsophyll / Bract
Protects floral bud
Bracts
📖
Parts of leaf

Table 1: Leaf Parts

Part
Synonym
Feature
Leaf base
Hypopodium
Basal part of leaf
Petiole
Mesopodium / leaf stalk
Develops last in leaf
Lamina
Epipodium / leaf blade
Supported by midrib, veins and veinlets
▢ Leaf Base:
  • •
    Pulvinous → swollen leaf base
  • •
    Sheathing leaf base → expanded base covering stem partially/completely; common in monocots, grasses, wheat, sugarcane, maize
▢ Petiole Types:

Table 1: Petiole Modifications

Type
Feature
Examples
Winged petiole
Petiole with wing-like expansion
Citrus, sweet pea
Tendrillar petiole
Petiole modified into tendril
Nepenthes, Clematis
Bulbous petiole
Spongy, inflated, aerenchymatous
Eichhornia
Phyllode
Green flattened leaf-like structure from petiole
Australian acacia, Parkinsonia
▢ Other Structures:
  • •
    Stipules → at base of leaf
  • •
    Stipels → at base of leaflets
📖
Venation
▢ Definition: Arrangement of veins and veinlets in lamina

Table 1: Types of Venation

Type
Common in
Subtypes / Exceptions
Reticulate venation
Dicots
Pinnate/unicostate; palmate/multicostate → convergent/divergent
Parallel venation
Monocots
Pinnate/unicostate; palmate/multicostate → convergent/divergent
Furcate venation
Forked venation
—
▢ Reticulate Venation Exceptions in Dicots:
  • •
    Calophyllum
  • •
    Corymbium
  • •
    Eryngium
▢ Parallel Venation Exceptions in Monocots:
  • •
    Alocasia
  • •
    Arisaema
  • •
    Colocasia
  • •
    Dioscorea
  • •
    Smilax
📖
Types of leaves

Table 1: Simple and Compound Leaves

Type
Subtypes
Simple leaf
Single lamina
Pinnately compound leaf
Unipinnate, bipinnate, tripinnate, decompound
Palmately compound leaf
Unifoliate, bifoliate, trifoliate, quadrifoliate, multifoliate/digitate
📚
INFLORESCENCE
▢ Introduction:
❖ Definition: Arrangement and mode of distribution of flowers on shoot system
❖ Purpose: Maximum pollination
❖ Anthotaxy: Arrangement of flowers in inflorescence
▢ Solitary Inflorescence:

Table 1: Solitary Flowers

Type
Feature
Example
Solitary terminal
Single flower at tip
Papaver somniferum / poppy
Solitary axillary
Single flower in leaf axil
Hibiscus rosa-sinensis / China rose
▢ Racemose vs Cymose:

Table 1: Racemose vs Cymose Inflorescence

Feature
Racemose
Cymose
Image 1
Growth
Indefinite / indeterminate / continuous
Definite / determinate / discontinuous
Peduncle
Does not end in flower
Ends in flower
Arrangement
Acropetal / centripetal
Basipetal / centrifugal
Flower age
Older at base, younger at apex
Older at top/centre, younger at base/periphery
▢ Simple Racemose:

Table 1: Simple Racemose Inflorescences

Type
Feature
Examples
Typical raceme
Unbranched peduncle with stalked/pedicellate flowers
Radish
Corymb
Lower flowers long stalked, upper flowers short stalked; all flowers at same level
Iberis amara / candytuft
Corymbose raceme
Mixture of raceme and corymb
Mustard
Spike
Sessile flowers in acropetal succession; most common type
Bottle brush / Callistemon
Catkin / Amentum
Unisexual, sessile, often hanging spike
Jackfruit, pineapple, mulberry, Salix, Betula
Umbel
Flowers arise from same point with equal pedicels; reduced peduncle; involucre present
Umbellifers
Head / Capitulum
Flattened peduncle forms receptacle with sessile florets; ray florets peripheral, disc florets central
Marigold, sunflower, chrysanthemum
Spadix
Thick fleshy peduncle; covered by spathe; male flowers top, neuter middle, female base
Colocasia, Arisaema, female maize
❖ Involucre Found In:
  • •
    Capitulum
  • •
    Cyathium
  • •
    Umbel
▢ Compound Racemose:

Table 1: Compound Racemose Inflorescences

Type
Feature
Examples
Panicle / compound raceme
Branched raceme
Cassia fistula, male maize
Compound corymb
Formed by small corymbs
Cauliflower
Compound umbel
Umbel of umbels
Coriander
Spikelets
Spike of spikes
Graminae, Amaranthus
Compound spadix
Branched spadix
Palm, coconut, banana
❖ Notes:
  • •
    Bracts of main corymb = involucre
  • •
    Bracts of small corymb = involucels
  • •
    In banana, whole inflorescence covered by spathe
▢ Cymose Inflorescence:

Table 1: Cymose Types

Type
Feature
Examples
Monochasial / uniparous cyme
Terminal branch ends in one flower and pushes one lateral branch
Helicoid, scorpoid
Helicoid cyme
Flowers borne on one side
Begonia, Drosera, Heliotropium
Scorpoid cyme
Flowers alternately on both sides; zig-zag axis
Freesia, Ranunculus, cotton
Rhipidium
Scorpoid cyme with all flowers in one plane
Solanum nigrum
Dichasial / biparous cyme
Terminal flower subtended by two lateral branches
Common cyme
Multichasial / polychasial cyme
Flowers borne on more than two sides
—
Scapigerous cyme umbel
Leafless scape from ground bears umbellate cyme covered by spathes
Onion
▢ Special Inflorescence:

Table 1: Special Inflorescences

Type
Feature
Examples / Fruit
Cyathium
Cup-shaped involucre; looks like single flower; one female flower surrounded by many male flowers; ratio 1:many or 1:5
Euphorbia
Hypanthodium
Fleshy pear/flask-shaped receptacle with hollow cavity
Ficus; develops into syconus fruit
Verticillaster
Biparous/dichasial cyme ending in uniparous cyme; condensed cyme
Labiatae: Ocimum, Salvia, mint
📚
FLOWER
▢ Introduction:

Table 1: Flower Basics

Feature
Point
Flower
Modified shoot with condensed internodes and floral parts as modified leaves
Function
Forms fruit and seed; helps reproduction
Cauliflory / trunkiflory
Flowers develop on old stem from dormant bud
Largest flower
Rafflesia
Smallest flower
Wolffia microscopica
Non-essential parts
Sepals, petals
Essential parts
Stamen, carpel
Pedicel
Stalk of flower
Peduncle
Stalk of inflorescence
Petiole
Stalk of leaf
Thalamus / torus
Swollen end of pedicel; 4 nodes and short internodes
▢ Flower Characters:

Table 1: General Flower Types

Type
Meaning
Examples
Complete flower
All 4 whorls present: calyx, corolla, androecium, gynoecium
—
Incomplete flower
One or more whorls absent
—
Bisexual / monoclinous / perfect / intersexual
Both androecium and gynoecium present
Mustard, hibiscus, Datura, pea, onion, grass, disc floret
Unisexual / diclinous
Only one sex present
Cucumber, ray floret
Pistillate
Only gynoecium
Ray floret
Staminate
Only androecium
Cucurbits, papaya, mulberry
Monoecious
Male and female flowers on same plant
Maize, castor, Colocasia, Ficus, cucurbits, Pinus, mango
Dioecious
Male and female flowers on separate plants
Cycas, papaya, mulberry, lapsi
Polygamous
Unisexual, bisexual and neuter flowers on same plant
Mango, Polygonum
▢ Flower Opening and Arrangement:

Table 1: Flower Conditions

Condition
Meaning
Examples
Cleistogamous
Flowers never open; do not expose pollen
Arachis, Oxalis, Viola, Mirabilis, balsam
Chasmogamous
Flowers open at maturity
Commelina benghalensis has both cleistogamy and chasmogamy
Cyclic
Floral parts arranged in circles
—
Acyclic / spiral
Floral parts arranged spirally; primitive angiosperm character
—
Bracteate
Flower covered by bract/involucre
Compositae, Euphorbiaceae, Gramineae
Ebracteate
Bract absent
Brassicaceae, Solanaceae
▢ Symmetry and Ovary Position:

Table 1: Symmetry and Floral Position

Type
Meaning
Examples
Actinomorphic
Radially symmetrical; divisible into two equal halves by any vertical plane
Cruciferae, Mimosaceae, Malvaceae, Liliaceae, disc floret
Zygomorphic
Bilaterally symmetrical; only one plane divides into two equal halves
Papilionaceae, Caesalpiniaceae, Graminae, ray floret
Slightly zygomorphic
Due to oblique ovary
Solanaceae
Hypogynous
Ovary superior
Brassicaceae
Perigynous
Ovary half-inferior / middle
Pea, bean, rose
Epigynous
Ovary inferior
Compositae, cucurbits
▢ Floral Whorl Condition:

Table 1: Calyx and Corolla Condition

Type
Meaning
Examples
Achlamydous
Calyx and corolla absent
Euphorbiaceae, Piperaceae
Monochlamydous / Haplochlamydous
Only one whorl present, either calyx or corolla
Polygonaceae, Gramineae, Liliaceae
Dichlamydous
Both calyx and corolla present
Most flowers
Perianth
Calyx and corolla not differentiated
Tepals
▢ Floral Internodes:

Table 1: Internodes of Flower

Structure
Internode
Position
Examples
Anthophore
1st internode
Between calyx and corolla
Silene
Androphore / gonophore
2nd internode
Between corolla and androecium
Passiflora
Gynophore
3rd internode
Between androecium and gynoecium
Gynandropsis, Capparis
Gynandrophore / androgynophore
Both androphore and gynophore distinct
—
Gynandropsis, Cleome
Carpophore
Raised thalamus between fused carpels
—
Coriander
▢ Dichogamy and Anthesis:

Table 1: Maturation Terms

Term
Meaning
Examples
Dichogamy
Pollen grains and ovules mature at different times
—
Protandry
Pollen grains mature earlier
Salvia, cotton, sunflower, rose, moss
Protogyny
Ovules / carpels mature earlier
Ficus, Magnolia, four o'clock plant
Anthesis
Opening of floral bud / time when flower is fully open and functional
—
▢ Calyx:

Table 1: Calyx / Sepal

Feature
Point
Position
Outermost floral whorl
Unit
Sepal
Colour
Usually green / sepaloid
Petaloid sepal
Sepal with colour other than green
Function
Protection of bud
Polysepalous
Sepals free
Gamosepalous
Sepals fused
Modified sepals
Pappus or scales in Compositae
Epicalyx
Additional whorl of sepal-like bracteoles; Malvaceae, Cucurbitaceae
❖ Sepal Persistence:

Table 1: Sepal Duration

Type
Feature
Examples
Caducous
Falls immediately after flower opens
Papaver
Deciduous
Withers after fertilization during fruit maturation
Cucurbitaceae, Malvaceae, Brassicaceae
Persistent
Does not fall after fertilization
Solanaceae
▢ Corolla:

Table 1: Corolla / Petal

Feature
Point
Unit
Petal
Colour
Various
Function
Attract insects for pollination
Polypetalous
Petals free
Gamopetalous
Petals fused
❖ Corolla Shapes:

Table 1: Corolla Forms

Shape
Feature
Example
Cruciform
Cross-like
Cruciferae
Papilionaceous
Butterfly-like
Pea family / Papilionaceae
Ligulate
Strap / tongue-like
Ray floret of Compositae
Tubular
Tube-like
Disc floret of Compositae, Solanaceae
Campanulate
Inverted bell-shaped
Solanaceae
Bilabiate
Two-lipped zygomorphic gamopetalous corolla
Ocimum
Personate
Masked bilabiate with narrow opening
Antirrhinum
Infundibuliform
Funnel-shaped
Datura
❖ Aestivation:

Table 1: Aestivation Types

Type
Feature
Examples
Valvate
Margins meet but do not overlap
Cruciferae, Mimosaceae, Asteraceae
Contorted / twisted
Margins overlap regularly
Malvaceae petals
Quincuncial
Two outer, two inner, one twisted
—
Descending imbricate / vexillary
Posterior petal overlaps lateral; laterals overlap anterior
Papilionaceae
Ascending imbricate
Posterior overlapped by lateral; lateral overlapped by anterior
Caesalpiniaceae
▢ Perianth:

Table 1: Perianth

Term
Meaning
Perianth
Indistinct calyx and corolla
Tepal
Unit of perianth
Polyphyllous
Tepals free
Gamophyllous
Tepals fused
Epiphyllous
Stamen fused with tepal; Liliaceae
Lodicule
Structure equivalent to perianth in grasses
▢ Androecium:
❖ Introduction:

Table 1: Androecium

Feature
Point
Nature
Male reproductive whorl
Unit
Stamen
Anther
Male sporophyll / fertile part
Connective
Vascular supply to anther
Filament
Stalk
Typical anther
Dithecous / bithecous and tetrasporangiate
Each anther lobe
2 pollen sacs
Monothecous anther
Single lobe and bisporangiate; Malvaceae
Monosporangiate anther
Arceuthobium
Pollinium
Pollen grains in sac-like structure; Asclepiadaceae, Orchidaceae
Staminode
Sterile undeveloped stamen
❖ Stamen Position and Fusion:

Table 1: Stamen Conditions

Condition
Meaning
Example
Introse
Stamens remain inside closed flower
Pisum sativum
Extrose
Stamens longer than petals and protrude
Malvaceae
Polyandrous
Stamens free
—
Episepalous
Stamen fused with sepals
Verbena
Epipetalous
Stamen fused with petals
Malvaceae
Epiphyllous
Stamen fused with tepals
Liliaceae
Gynandrous
Stamen fused with carpel forming gynostegium
Calotropis
❖ Anther Fixation:

Table 1: Anther Fixation

Type
Feature
Example
Basifixed
Filament attached at base of anther
Brassicaceae, Solanaceae
Dorsifixed
Filament attached dorsally
Legumes
Adnate
Filament attached along whole length of anther
Magnoliaceae
Versatile
Anther attached near middle of connective and freely movable
Graminae
❖ Cohesion of Stamens:

Table 1: Cohesion of Stamens

Type
Fusion
Example
Synandrous
Both anther and filament fused
Cucurbitaceae
Syngenesious
Anthers fused, filaments free
Compositae / Asteraceae
Monoadelphous
Filaments fused into one bundle; anthers free
Malvaceae
Diadelphous
Filaments fused into two bundles; anthers free
Papilionaceae
Polyadelphous
Filaments fused into more than two bundles; anthers free
Citrus, castor, Bombax
❖ Length of Stamens:

Table 1: Stamen Length

Type
Feature
Example
Didynamous
4 stamens; outer 2 long, inner 2 short
Labiatae / Ocimum
Tetradynamous
6 stamens; outer 2 short, inner 4 long
Brassicaceae
Heterostemony
Stamens of different lengths
Caesalpiniaceae
▢ Gynoecium:
❖ Introduction:

Table 1: Gynoecium / Carpel

Term
Meaning
Gynoecium
Female reproductive whorl
Unit
Carpel
Carpel parts
Stigma, style, ovary
Pistil
Single carpel or fused group of carpels
Simple gynoecium
One carpel / monocarpellary
Compound gynoecium
Two or more carpels
Apocarpous
Carpels free
Syncarpous
Carpels fused
❖ Carpel vs Pistil:

Table 1: Carpel vs Pistil

Carpel
Pistil
Female unit consisting of stigma, style and ovary
May be one carpel or fused collection of carpels
Number counted by separate carpels
Number counted by separate ovaries
Can be free or fused
Represents functional female reproductive structure
❖ Style and Stigma:

Table 1: Style and Stigma

Feature
Type / Example
Terminal style
Arises from top of ovary; Brassicaceae, Solanaceae
Lateral style
Arises from lateral side of ovary; Graminae
Gynobasic style
Arises from base of ovary; Labiatae e.g. Salvia, Ocimum
Bifid stigma
Compositae, Solanaceae, Brassicaceae
Trifid stigma
Liliaceae
Tetrafid stigma
Labiatae
Pentafid stigma
Malvaceae
❖ Ovary Locules:

Table 1: Types of Ovary by Locule

Type
No. of chambers
Unilocular
1
Bilocular
2
Trilocular
3
Tetralocular
4
Pentalocular
5
Multilocular
Many
❖ Placentation:
◉ Definition: Arrangement of ovules in ovary

Table 1: Types of Placentation

Type
Mnemonic
Examples / Family
Parietal
PCC
Cruciferae, Cucurbitaceae
Axile
SoMaLiA
Solanaceae, Malvaceae, Liliaceae
Marginal
MaLe
Leguminosae / Papilionaceae
Basal
BAG
Asteraceae, Graminae
❖ Gynoecium by Carpel Number:

Table 1: Types of Gynoecium

Type
Number of carpels
Simple gynoecium / monocarpellary
1
Bicarpellary
2
Polycarpellary
Many
📚
HIGH-YIELD RECALL

Table 1: Morphology One-Liners

Fact
Answer
Study of form and structure
Morphology
Study of flowering plants / flowers
Anthology
First flowering plant appeared
Jurassic period
Leaf arrangement
Phyllotaxy
Young leaf arrangement in bud
Prefoliation
Individual young leaf arrangement
Ptyxis
Circinate ptyxis
Fern, Cycas, banana
Inflorescence flower arrangement
Anthotaxy
Flower stalk
Pedicel
Inflorescence stalk
Peduncle
Leaf stalk
Petiole
Swollen end of pedicel
Thalamus / torus
Root origin
Radicle
Lateral roots arise from
Pericycle
Root origin nature
Endogenous
Root lacks
Nodes, internodes, leaves, flowers
Nodulated roots
Legumes
Root nodule bacterium
Rhizobium
Pigment in root nodules
Leghaemoglobin
Breathing roots
Pneumatophores
Pneumatophore pores
Pneumatothodes
Mycorrhizal root lacks
Root cap and root hairs
Prop root
Ficus benghalensis
Stilt root
Sugarcane, maize
Velamen tissue
Epiphytic orchid root
Sweet potato root
Tuberous adventitious root
Stem origin
Plumule
Stem hairs
Usually multicellular
Stem branches arise
Exogenously
Culm
Grass family
Excurrent stem
Conifers
Pseudostem
Banana
Caudex
Cycas, coconut
Thorn
Modified stem / axillary bud
Spine
Modified leaf
Prickle
Superficial outgrowth
Lianas
Woody tropical climbers
Runner
Oxalis, Cynodon
Offset
Pistia, Eichhornia
Sucker
Banana, mint
Stolon
Strawberry
Bulb
Onion, garlic
Corm
Colocasia, Amorphophallus
Rhizome
Ginger, turmeric
Tuber
Potato
Phylloclade
Opuntia
Cladode
Asparagus
First leaf
Prophyll
All leaf types in plant
Phyllome
Single-leaf plant
Monophyllaea
Two-leaf plant
Welwitschia
Dicot leaf
Dorsiventral
Monocot leaf
Isobilateral
Petiole develops
Last
Bulbous petiole
Eichhornia
Phyllode
Australian acacia
Reticulate venation
Common in dicots
Parallel venation
Common in monocots
Racemose growth
Indeterminate
Cymose growth
Determinate
Most common inflorescence
Spike
Capitulum
Sunflower
Spadix
Colocasia, Arisaema
Cyathium
Euphorbia
Hypanthodium
Ficus
Verticillaster
Labiatae
Largest flower
Rafflesia
Smallest flower
Wolffia microscopica
Flower is
Modified shoot
Complete flower
Calyx + corolla + androecium + gynoecium
Bisexual flower
Androecium + gynoecium present
Cauliflory
Flower on old stem
Cleistogamy
Closed flower
Actinomorphic
Radially symmetrical
Zygomorphic
Bilaterally symmetrical
Hypogynous
Superior ovary
Epigynous
Inferior ovary
Calyx unit
Sepal
Corolla unit
Petal
Perianth unit
Tepal
Valvate aestivation
Margins meet, no overlap
Vexillary aestivation
Papilionaceae
Typical anther
Dithecous + tetrasporangiate
Monothecous anther
Malvaceae
Pollinium
Calotropis, orchids
Syngenesious stamens
Asteraceae
Monoadelphous stamens
Malvaceae
Diadelphous stamens
Papilionaceae
Tetradynamous stamens
Brassicaceae
Gynobasic style
Labiatae
Marginal placentation
Leguminosae
Axile placentation
Solanaceae, Malvaceae, Liliaceae
Parietal placentation
Cruciferae, Cucurbitaceae
Basal placentation
Asteraceae, Graminae
Q1.
Phylloclade and cladode are morphological adaptations for photosynthesis and water storage, found in
📅CEE MBBS 2025
Q2.
In which inflorescence does the main axis show definite growth and terminate in a flower?
📅CEE MBBS 2025
Q3.
A fruit formed from many free carpels of a single flower is:
📅CEE BDS 2024
Q4.
A phylloclade is a modified:
📅CEE BDS 2024
Q5.
Bicollateral vascular bundles commonly occur in:
📅CEE BDS 2024
Q6.
Stilt roots of sugarcane originate from the:
📅CEE MBBS 2023
Q7.
Phylloclades and cladodes are common adaptations of:
📅CEE BDS 2023
Q8.
A leaf lamina is typically attached to the stem through the:
📅CEE BDS 2023
Q9.
Which of the following is a fleshy berry in which the fruit wall (pericarp) forms much of the edible portion?
📅CEE BDS 2023
Q10.
The edible white flesh of coconut is:
📅CEE MBBS 2022
Q11.
The arrangement of petals in pea flowers is:
📅CEE MBBS 2022
Q12.
The inflorescence of sunflower is:
📅CEE MBBS 2021
Q13.
Which is an underground stem modification for storage and perennation?
📅CEE BDS 2021
Q14.
The edible fleshy part of litchi is the:
📅CEE BDS 2021
Q15.
An apocarpous gynoecium has:
📅CEE BDS 2021