19Plant Movement

📚
MOVEMENT
Definition: Living plants → positional change due to environmental/endogenous causes
Cause: Sensitivity / irritability of protoplasm
Level:
  • Whole organism
  • Organ level
  • Cellular level
Cellular Example: Cyclosis = cytoplasmic streaming; cytoplasm flows like river inside living cell
📚
STIMULUS & RESPONSE
Stimulus: External environmental change → reaction in plant/organ
Threshold Intensity: Minimum stimulus intensity → response induction
Response: Visible change/reaction of plant/organ due to stimulus
Reaction Time / Presentation Time: Interval: stimulus presentation → actual response development
Stimulus-Based Movement:

Table 1: Nature of Stimulus

Type
Cause
Also Called
Autonomous
Internal stimulus
Spontaneous
Paratonic
External stimulus
Induced
Stimuli Prefixes:

Table 1: Stimulus → Prefix

Stimulus
Prefix
Light
Photo
Temperature
Thermo
Water
Hydro
Chemical
Chemo
Water current
Rheo
Gravity
Geo
Touch / Contact
Thigmo / Hapto
Shock
Seismo
📚
HYGROSCOPIC MOVEMENT
Occurrence:
  • Bryophytes
  • Pteridophytes
Examples:
  • Twisting movement of elater
  • Sporangial wall of fern
📚
TYPES OF MOVEMENT

Table 1: Major Types

Feature
Movement of Locomotion
Movement of Curvature
Common In
Lower plants
Higher plants
Change
Position of whole body
Position of body parts only
Main Types
Autonomic + Paratonic
Movement of variation + Movement of growth
📖
Movement of locomotion
Autonomic / Self Movement:

Table 1: Autonomic Locomotory Movements

Type
Mechanism
Examples
Amoeboid
Pseudopodia-like structure
Synchytrium / slime mould
Ciliary
Cilia / flagella
Chlamydomonas, Volvox
Cyclosis / Streaming
Protoplasmic streaming
Chloroplast movement in Elodea
Cyclosis Types:

Table 1: Cyclosis

Type
Movement Direction
Example
Rotation
Clockwise OR anticlockwise; single direction
Leaf of Hydrilla, Vallisneria
Circular
Clockwise + anticlockwise; all possible directions
Staminal hairs of Tradescantia
Paratonic / Induced Movement:
Name: Tactic / Taxis movement

Table 1: Tactic Movements

Type
Stimulus
Key Points
Examples
Phototactic
Light
Movement: dark → diffused light; intense light ↔ diffused light
Zoospores of Chlamydomonas, Volvox
Thermotactic
Temperature
Cold water ↔ warm water
Chlamydomonas, bacteria
Chemotactic
Chemical, e.g., malic acid
Antherozoid movement: antheridium → archegonium
Bryophytes, pteridophytes
Rheotactic
Water current
Movement due to water current
Aquatic algae
📖
Movement of curvature
Movement of Variation:
  • Change in position without growth
  • Mostly turgor/osmotic change
Movement of Growth:
  • Change in position due to growth
  • Growth differential → curvature
📝
Movement of Variation
Autonomic:
Example: Leaves of Desmodium gyrans = Indian telegraph plant
Key Point: Lateral leaflets → continuous up-down movement
Paratonic / Induced / Nastic Movement:

Table 1: Nastic Movements

Type
Stimulus
Key Point
Examples
Photonasty
Light
Response to light intensity
Head of sunflower → helionasty
Thermonasty
Temperature
Flowering/opening at higher temperature
Tulip, crocus flower
Nyctinasty / Sleep movement
Light + temperature
↓ light intensity → sleep movement of leaves
Oxalis + Albizzia: open day, close night; Oenothera: open night, close day
Seismonasty / Osmotic / Turgor movement
Shock / touch
Water loss from pulvinus + pulvinules → drooping
Leaves of Mimosa pudica
Thigmonasty / Haptonasty
Touch / contact
Touch-induced movement
Insectivorous plants; hairs of Drosera
📝
Movement of Growth
Autonomic Growth Movement:

Table 1: Autonomic Growth Movements

Type
Growth Pattern
Movement
Examples
Hyponasty
More growth of lower surface / abaxial side
Part moves upward
Closed flower petal; young fern leaf / circinately coiled leaf
Epinasty
More growth of upper surface / adaxial side
Part moves downward
Opening of flower; erect fern leaf / mature frond
Nutational movement
Alternate growth on either surface
Zig-zag / forward curvature
Zig-zag shoot apex; forward growth of branches
Hyponasty + epinasty → common in bifacial structures: leaf, sepal, petal
Paratonic Growth Movement:
Image 1
Also Called: Tropic movement; process = tropism

Table 1: Tropic Movements

Type
Stimulus
Positive Response
Negative Response
Examples / Notes
Phototropic
Light
Shoots
Roots
Differential hormonal effect; auxin accumulates on shaded side; coleoptile bends toward light due to greater elongation on shaded side
Geotropic
Gravity
Roots
Shoots
Main root = +ve geotropic; main shoot = -ve geotropic
Hydrotropic
Water
Roots
Shoots
Important for young roots
Thigmotropic / Haptotropic
Touch / contact
Contact side response
Tendrils bending on support; haustoria in Cuscuta
Chemotropic
Chemical, e.g., malic acid
Growth toward chemical source
Pollen tube growth toward embryo sac
📄
Geotropism Types

Table 1: Types of Geotropic Growth

Type
Growth Direction vs Gravity
Examples
Orthogeotropic
Parallel to gravity
Main roots → +ve orthogeotropic; main shoot → -ve orthogeotropic
Diageotropic
Right angle to gravity
Tertiary root; rhizome; prostrate plants; leaves
Plagiogeotropic
Between \(0-90^\circ\) / \(90-180^\circ\)
Secondary/lateral roots → +ve plagiogeotropic; stem branches → -ve plagiogeotropic
Apogeotropic / Ageotropic
No effect of gravity
Respiratory roots/pneumatophores of mangroves; coralloid root of Cycas
📚
INSTRUMENTS & SPECIAL TROPISMS
Clinostat: Instrument → eliminates effect of gravity on plant
Traumatotropism: Movement induced by injury
📚
READ & DIGEST
Bulliform Cells:
  • Grass leaves → folding + unfolding
  • Specialized bulliform cells
  • Bulliform cells → turgor movements
Nastic vs Tropic: Nastic movements = non-directional; tropic movements = directional
Scototropic Movement: Induced by night
Xerochasy: Peristome teeth of moss
Earthworm: Moves down soil → +ve geotropism
Telotaxis: Honey bee drawn toward flower