18Plant Movement

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MOVEMENT
▢ Definition: Living plants → positional change due to environmental/endogenous causes
▢ Cause: Sensitivity / irritability of protoplasm
▢ Level:
  • •
    Whole organism
  • •
    Organ level
  • •
    Cellular level
▢ Cellular Example: Cyclosis = cytoplasmic streaming; cytoplasm flows like river inside living cell
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STIMULUS & RESPONSE
▢ Stimulus: External environmental change → reaction in plant/organ
▢ Threshold Intensity: Minimum stimulus intensity → response induction
▢ Response: Visible change/reaction of plant/organ due to stimulus
▢ Reaction Time / Presentation Time: Interval: stimulus presentation → actual response development
▢ Stimulus-Based Movement:

Table 1: Nature of Stimulus

Type
Cause
Also Called
Autonomous
Internal stimulus
Spontaneous
Paratonic
External stimulus
Induced
▢ Stimuli Prefixes:

Table 1: Stimulus → Prefix

Stimulus
Prefix
Light
Photo
Temperature
Thermo
Water
Hydro
Chemical
Chemo
Water current
Rheo
Gravity
Geo
Touch / Contact
Thigmo / Hapto
Shock
Seismo
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HYGROSCOPIC MOVEMENT
▢ Occurrence:
  • •
    Bryophytes
  • •
    Pteridophytes
▢ Examples:
  • •
    Twisting movement of elater
  • •
    Sporangial wall of fern
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TYPES OF MOVEMENT

Table 1: Major Types

Feature
Movement of Locomotion
Movement of Curvature
Common In
Lower plants
Higher plants
Change
Position of whole body
Position of body parts only
Main Types
Autonomic + Paratonic
Movement of variation + Movement of growth
📖
Movement of locomotion
▢ Autonomic / Self Movement:

Table 1: Autonomic Locomotory Movements

Type
Mechanism
Examples
Amoeboid
Pseudopodia-like structure
Synchytrium / slime mould
Ciliary
Cilia / flagella
Chlamydomonas, Volvox
Cyclosis / Streaming
Protoplasmic streaming
Chloroplast movement in Elodea
▢ Cyclosis Types:

Table 1: Cyclosis

Type
Movement Direction
Example
Rotation
Clockwise OR anticlockwise; single direction
Leaf of Hydrilla, Vallisneria
Circular
Clockwise + anticlockwise; all possible directions
Staminal hairs of Tradescantia
▢ Paratonic / Induced Movement:
❖ Name: Tactic / Taxis movement

Table 1: Tactic Movements

Type
Stimulus
Key Points
Examples
Phototactic
Light
Movement: dark → diffused light; intense light ↔ diffused light
Zoospores of Chlamydomonas, Volvox
Thermotactic
Temperature
Cold water ↔ warm water
Chlamydomonas, bacteria
Chemotactic
Chemical, e.g., malic acid
Antherozoid movement: antheridium → archegonium
Bryophytes, pteridophytes
Rheotactic
Water current
Movement due to water current
Aquatic algae
📖
Movement of curvature
▢ Movement of Variation:
  • •
    Change in position without growth
  • •
    Mostly turgor/osmotic change
▢ Movement of Growth:
  • •
    Change in position due to growth
  • •
    Growth differential → curvature
📝
Movement of Variation
▢ Autonomic:
❖ Example: Leaves of Desmodium gyrans = Indian telegraph plant
❖ Key Point: Lateral leaflets → continuous up-down movement
▢ Paratonic / Induced / Nastic Movement:

Table 1: Nastic Movements

Type
Stimulus
Key Point
Examples
Photonasty
Light
Response to light intensity
Head of sunflower → helionasty
Thermonasty
Temperature
Flowering/opening at higher temperature
Tulip, crocus flower
Nyctinasty / Sleep movement
Light + temperature
↓ light intensity → sleep movement of leaves
Oxalis + Albizzia: open day, close night; Oenothera: open night, close day
Seismonasty / Osmotic / Turgor movement
Shock / touch
Water loss from pulvinus + pulvinules → drooping
Leaves of Mimosa pudica
Thigmonasty / Haptonasty
Touch / contact
Touch-induced movement
Insectivorous plants; hairs of Drosera
📝
Movement of Growth
▢ Autonomic Growth Movement:

Table 1: Autonomic Growth Movements

Type
Growth Pattern
Movement
Examples
Hyponasty
More growth of lower surface / abaxial side
Part moves upward
Closed flower petal; young fern leaf / circinately coiled leaf
Epinasty
More growth of upper surface / adaxial side
Part moves downward
Opening of flower; erect fern leaf / mature frond
Nutational movement
Alternate growth on either surface
Zig-zag / forward curvature
Zig-zag shoot apex; forward growth of branches
Hyponasty + epinasty → common in bifacial structures: leaf, sepal, petal
▢ Paratonic Growth Movement:
Image 1
❖ Also Called: Tropic movement; process = tropism

Table 1: Tropic Movements

Type
Stimulus
Positive Response
Negative Response
Examples / Notes
Phototropic
Light
Shoots
Roots
Differential hormonal effect; auxin accumulates on shaded side; coleoptile bends toward light due to greater elongation on shaded side
Geotropic
Gravity
Roots
Shoots
Main root = +ve geotropic; main shoot = -ve geotropic
Hydrotropic
Water
Roots
Shoots
Important for young roots
Thigmotropic / Haptotropic
Touch / contact
Contact side response
Tendrils bending on support; haustoria in Cuscuta
Chemotropic
Chemical, e.g., malic acid
Growth toward chemical source
Pollen tube growth toward embryo sac
📄
Geotropism Types

Table 1: Types of Geotropic Growth

Type
Growth Direction vs Gravity
Examples
Orthogeotropic
Parallel to gravity
Main roots → +ve orthogeotropic; main shoot → -ve orthogeotropic
Diageotropic
Right angle to gravity
Tertiary root; rhizome; prostrate plants; leaves
Plagiogeotropic
Between \(0-90^\circ\) / \(90-180^\circ\)
Secondary/lateral roots → +ve plagiogeotropic; stem branches → -ve plagiogeotropic
Apogeotropic / Ageotropic
No effect of gravity
Respiratory roots/pneumatophores of mangroves; coralloid root of Cycas
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INSTRUMENTS & SPECIAL TROPISMS
▢ Clinostat: Instrument → eliminates effect of gravity on plant
▢ Traumatotropism: Movement induced by injury
📚
READ & DIGEST
▢ Bulliform Cells:
  • •
    Grass leaves → folding + unfolding
  • •
    Specialized bulliform cells
  • •
    Bulliform cells → turgor movements
▢ Nastic vs Tropic: Nastic movements = non-directional; tropic movements = directional
▢ Scototropic Movement: Induced by night
▢ Xerochasy: Peristome teeth of moss
▢ Earthworm: Moves down soil → +ve geotropism
▢ Telotaxis: Honey bee drawn toward flower
Q1.
Opening of flowers and drooping of buds in classical plant movement terminology are commonly associated with:
📅CEE MBBS 2023
Q2.
Downward bending of plant organs due to greater growth on the upper surface is:
📅CEE MBBS 2022
Q3.
The instrument used to cancel the directional stimulus of gravity in experiments is:
📅IOM 2014
Q4.
A plant detects the direction of light using:
📅IE 2007
Q5.
Short-day plants flower when they experience:
Q6.
The phenomenon in which a short light pulse interrupts a long dark period is called:
📅BPKIHS 2004
Q7.
Which instrument minimizes directional gravity effects by slowly rotating a plant?
📅IOM 2014
Q8.
Plants respond to directional blue light through the action of:
📅IE 2007